Machilus japonica var. kusanoi is an endemic variety to Taiwan. Bisexual flowers are produced from January to March. Inflorescence primordia occur at shoot apices in early January. Anther primordia are formed by mid-January, and composed of a protoderm and a central cell mass. The anther wall layers consist of the epidermis, the one-layered endothecium, two middle layers and the tapetum of one layer. The tapetal cells are amoeboid type of characteristics. Pollen mother cells undergo meiosis through successive cytokinesis and give rise to tetrads of microspores, which are mostly isobilaterally arranged. Mature pollen grains are 2-celled,globose and 30~45μm in diameter. Anthers are dehiscent by valves, which contain endothecial cells with uneven-thickened wall.