台灣穿山甲（Manis pentadactyla pentadactyla）為中華穿山甲的亞種，為穴居型夜行性動物，其食性專一，以螞蟻和白蟻為主食，但對於蟻種則較無偏好。為瞭解穿山甲掘洞的數量是否受地表螞蟻活動量影響。本研究在雙流森林遊樂區的帽子山步道與壽卡林道，各選取10個樣點，調查穿山甲舊有洞穴數量並記錄月新增洞穴，以及採集地表螞蟻。結果共獲得856隻螞蟻個體，分屬8亞科，22屬，36種，其中有11種僅採集到1隻個體。研究期間一共尋獲284個新增的穿山甲洞穴，且濕季（5-10月)新增的數量略高於乾季（11-4月)，有別於前人發現冬、春（乾季）兩季洞穴數量較多的現象。推測是因雙流地區的月均溫差異不大，對於地表螞蟻的數量影響較小；但乾、濕季分明，當雨量過大時會讓螞蟻數量短暫減少，穿山甲為了搜尋螞蟻而掘洞，才使濕季的洞穴數量提高。儘管本研究穿山甲每月新增洞穴數量與地表螞蟻數量沒有顯著相關性，但地表螞蟻數量較多的樣線會發現較多的穿山甲洞穴，且穿山甲對覓食地點具有高度忠誠度，顯示食物資源的多寡可能是影響穿山甲活動範圍分布的主要原因。

Taiwan pangolin (Mains pentadactyla pentadactyla) is a subspecies of Chinese pangolin, which is a burrowing nocturnal animal. It has a specific diet and mainly feeds on ants and termite, but it has not preference for ant species or termite species. In previous studies, it was observed that the new seasons of pangolin caves are mainly distributed in winter and spring (dry season). The number of ants species found in feces is significantly lower in summer compared to other seasons. To know whether the number of pangolin digging burrows is affected by the amount of surface ant activity, this study investigated the number of old caves in pangolins and recorded the number of new caves in the month and the activity of ants on the ground at the Mt. Maozi Trail and Shouka in the Shuangliu Nation Forest Recreation Area. During 17-month study period (Oct., 2018- Feb., 2020), A total of 36 species with 856 individuals in 22 genera belonging to 8 subfamilies were collected and identified. There were 284 burrows found during the study period. Formosa pangolins have dug burrows more in wet season (May-Oct.) than in dry season(Nov.-Apr.). It is speculated that the monthly average temperature difference in Shuangliu area is not obvious, but the dry and wet seasons are distinct, so compared with the temperature, the rainfall has a greater impact on the number of surface ants. When the rain is too large, it will temporarily reduce the number of ants and the pangolin dig more burrows. There is no significant correlation between the number of burrows and ants in the Shuangliu area, but the sample line with more surface ants will find more burrows. The pangolin also has a high degree of loyalty to foraging sites, indicating that the amount of food resources may be the main reason for affecting the distribution of pangolin activities.

林宗岐、吳文哲(2003)台灣螞蟻相 (膜翅目：蟻科) ─ 並附亞科與屬檢索表。國立台灣博物館年刊 46: 5-69。
林敬勛(2010)台東鸞山地區台灣穿山甲（Manis pentadactyla pentadactyla）活動範圍與洞穴利用之研究。碩士論文， 國立屏東科技大學野生動物保育研究所。68頁。
姜博仁、蔡哲民、蔡世超、吳禎祺、鄭薏如(2015) 錄音技術應用於野生動物調查之應用與評估。台灣林業季刊 41(4):33-38。
范中衍 (2005) 翡翠水庫台灣穿山甲洞穴棲地研究。碩士論文，國立臺灣大學森林學研究所。56頁。
梁竣傑(2017)台東鸞山地區土壤環境與穿山甲排遺中的白蟻物種組成。碩士論文， 國立屏東科技大學野生動物保育研究所。50頁。
許峰銓(2015) 蓮華池森林動態樣區垂直結構之螞蟻群聚研究。碩士論文，國立彰化師範大學生物學系碩士班。80頁。
陸聲山 (2005) 台灣北部山區穿山甲分布、現況及生態之研究（2/2）。農委會林業試驗所九十四年度科技計畫研究報告。2頁。
楚南仁博 (1941) 穿山甲の食物調查。科學的台灣 9 (3): 9-11。
董景生、山馥嫻、林宗岐(2009) 福山地區螞蟻監測及鑑定指南。行政院農業委員會農業試驗所。165頁。
劉啟斌(2003)恆春半島山地植群之分類。碩士論文，國立屏東科技大學森林系碩士班。152頁。
鍾富雅(2010) 蓮華池試驗林地棲與樹棲螞蟻群聚結構之研究。碩士論文，國立彰化師範大學生物學系碩士班。98頁。
Adam V., H. Jaroslav and J. Frouz. (2009) Sampling for ants in different-aged spruce forests: a comparison of methods. European Journal of Soil Biology 45(4): 301-305.
Andersen AN, Sparling GP (1997) Ants as indicators of restoration success: relationship with soil microbial biomass in the Australian seasonal tropics. Restor Ecology 5: 109–114.
Andersen, A. N., and J. D. Majer. (2004) Ants show the way down under: invertebrates as bioindicators in land management. Frontiers in Ecology and the Environment 2: 291-298.
Andersen, A. N. (1983) Species diversity and temporal distribution of ants in the semi-arid mallee region of north-western Victoria. Aust. J. Ecology 8: 127-137.
Andersen, A. N. (1986) Diversity, seasonality and community organization of ants at adjacent heath and woodland sites in South-Eastern Australia. Aust J. Zool. 34: 53-64.
Andersen, A. N. (1995) A classification of Australian ant communities, based on functional groups which parallel plant lifeforms in relation to stress and disturbance. J Biogeogr 22: 15–29.
Biesinger, Z. Y., & Haefner, J. W. (2005) Proximate cues for predator searching: a quantitative analysis of hunger and encounter rate in the ladybird beetle, Coccinella septempunctata. Animal Behaviour 69(1): 235-244.
Boakye, M. K., D. W. Pietersen, A. Kotze, D. L. Dalton, and R. Jansen (2015) Knowledge and uses of African pangolins as a source of traditional medicine in Ghana. Plos One 10: 117-199.
Boonzaaier, C., McGeoch, M. A., & Parr, C. L. (2007) Fine-scale temporal and spatial dynamics of epigaeic ants in Fynbos: sampling implications. African Entomology 15(1): 1-11.
Borgelt, A., & New, T. R. (2006) Pitfall trapping for ants (Hymenoptera, Formicidae) in mesic Australia: what is the best trapping period. Journal of Insect Conservation 10(1): 75-77.
Botes A, McGeoch MA, Robertson HG, Van Niekerk A, Davids HP, Chown SL (2006) Ants, altitude and change in the northern Cape Floristic Region. J Biogeogr 33: 71–90.
Brener, A. G. F., & Ruggiero, A. (1994) Leaf-cutting ants (Atta and Acromyrmex) inhabiting Argentina: patterns in species richness and geographical range sizes. Journal of Biogeography 391-399.
Bronikowski, A. M., & Altmann, J. (1996) Foraging in a variable environment: weather patterns and the behavioral ecology of baboons. Behavioral Ecology and Sociobiology 39(1): 11-25.
Chao, J. T. (1989) Studies on the conservation of the Formosan pangolin (Manis pentadactyla pentacactyla) In: General biology and current status. Council of Agriculture, Taipei, Taiwan.
Ching-Min Sun, Jai-Chyi Pei, Chung-Chi Lin, Hou-Feng Li, Chun-Chieh Liang (2017) The Feeding Ecology of Formosan Pangolin (Manis pendatactyla pendatactyla) in the Southern Coastal Mountain Range, Taiwan. In: 2017 13th International conference on One Plan Approach Conservation Planning and Formosan Pangolin PHVA Workshop. December 4-8, 2017. Taipei Zoo, Taiwan.
Clay, R. E., & Schneider, K. E. (2000) The ant (Hymenoptera: Formicidae) fauna of coastal heath in south-west Victoria: effects of dominance by Acacia sophorae and management actions to control it. Pacific Conservation Biology 6(2): 144-151.
Coll, M., & Guershon, M. (2002) Omnivory in terrestrial arthropods: mixing plant and prey diets. Annual review of entomology 47(1): 267-297.
Coulson, I. (1989) The pangolin (Manistemmincki Smuts, 1835) in Zimbabwe. African Journal of Ecology 27: 149-155.
Daniel J. Ingram, Daniel Willcox, Daniel W.S. Challender (2019) Evaluation of the application of methods used to detect and monitor selected mammalian taxa to pangolin monitoring. Global Ecology and Conservation 18: 1-25.
Daniel Willcox, Helen C. Nash , Scott Trageser, Hyeon Jeong Kim ,Lisa Hywood , Ellen Connelly , Godwill Ichu Ichu, Jonas KambaleNyumu , Carla Louise Mousset Moumbolou , Daniel J. Ingram,Daniel W.S. Challender (2019) Evaluating methods for detecting and monitoring pangolin(Pholidata: Manidae) populations. Global Ecology and Conservation 17: 1-25.
Bestelmeyer, B. T., Agosti, D., Alonso, L. E., Brandão, C. R. F., Brown, W. L., Delabie, J. H., & Silvestre, R. (2000) Field techniques for the study of ground-dwelling ant: an overview, description, and evaluation.
Donnelly, D. & Giliomee, J. H. (1985) Community structure of epigaeic ants (Hymenoptera: Formicidae) in fynbos vegetation in the Jonkershoek Valley. J. entomol. Soc. South Afr. 48: 248-257.
Dunn RR, Agosti D, Andersen AN, Arnan X, Bruhl CA, Cerda´ X, Ellison AM, Fisher BL, Fitzpatrick MC, Gibb H, Gotelli NJ, Gove AD, Guenard B, Janda M, Kaspari M, Laurent EJ, Lessard JP, Longino JT, Majer JD, Menke SB, McGlynn TP, Parr CL, Philpott SM, Pfeiffer M, Retana J, Suarez AV, Vasconcelos HL, Weiser MD, Sanders NJ (2009) Climatic drivers of hemispheric asymmetry in global patterns of ant species richness. Ecology Lett 12: 324–333.
Greenslade, P. J. M. (1964) Pitfall trapping as method for studying populations of Carabidae (Coleoptera). J. Anim. Ecology 33: 301-310.
Greenslade, P. J. M. & Greenslade, P. (1984) Soil surface insects of the Australian arid zone. Proceedings of the Sesquicentennial conference on arid Australia. Sydney: Australian Museum.
Hölldobler, B., and Wilson, E. O. (1990) The ants. Harvard University Press.
Holling, C. S. (1961) Principles of insect predation. Annual review of entomology 6(1): 163-182.
Huey, R. B., & Pianka, E. R. (1981) Ecological consequences of foraging mode. Ecology 62(4): 991-999.
Irshad, N., T. Mahmood, R. Hussain, and M. S. Nadeem (2015) Distribution, abundance and diet of the Indian pangolin (Manis crassicaudata). Animal Biology 65: 57-71.
Kaspari, M. (2000) Do imported fire ants impact canopy arthropods? Evidence from simple arboreal pitfall traps. The southwestern Naturalist 45: 118-122.
Koen, J. H. & Breytenbach, W. (1988) Ant species richness of fynbos and forest ecosystems in the southern cape. S. Afr. J. Zool. 23: 184-188.
Lahkar, D., Ahmed, M.F., Begum, R.H., Das, S.K., Lahkar, B.P., Sarma, H.K., Harihar, A. (2018) Camera-trapping survey to assess diversity, distribution and photographic capture rate of terrestrial mammals in the aftermath of the ethnopolitical conflict in Manas National Park, Assam, India. J. Threat Taxa 10: 12008-12017.
Lim, N. T., & Ng, P. K. (2008) Home range, activity cycle and natal den usage of a female Sunda pangolin Manis javanica (Mammalia: Pholidota) in Singapore. Endangered Species Research 4(1-2): 233-240.
Lopes, C. T. and H. L. Vasconcelos. (2008) Evaluation of Three Methods for Sampling Ground-Dwelling Ants in the Brazilian Cerrado. Neotropical Entomology 37: 399-405.
Luff, M. L. (1975) Some features influencing the efficiency of pit fall traps. Oecologia 19: 345-357.
Mahmood, T., K. Jabeen, I. Hussain, and A. R. Kayani (2013) Plant species association, burrow characteristics and the diet of the Indian pangolin, Manis crassicaudata, in the Potohar Plateau, Pakistan. Pakistan Journal of Zoology 45: 1533-1539.
Majer, J. D. (1983) Ants: bio-indicators of minesite rehabilitation, land-use, and land conservation. Environmental management 7(4): 375-383.
McDonough, C. M., M. J. Delaney, P. Q. Le, M. S. Blackmore & W. J. Loughry (2000) Burrow characteristics and habitat associations of armadillos in Brazil and the United States of America. Revista de Biologia Tropical 48(1): 109-120.
Mitrovich MJ, Matsuda T, Pease KH, Fisher RN (2010) Ants as a measure of effectiveness of habitat conservation planning in Southern California. Conserv Biology 24: 1239–1248.
Moo, S.S.B., Froese, G.Z.L., Gray, T.N.E.(2017) First structured camera-trap surveys in Karen State, Myanmar, reveal high diversity of globally threatened mammals. Oryx 52(3): 537-543.
Munyai, T. C., & Foord, S. H. (2012) Ants on a mountain: spatial, environmental and habitat associations along an altitudinal transect in a centre of endemism. Journal of Insect Conservation 16(5): 677-695.
Parr CL, Robertson HG, Biggs HC, Chown SL (2004) Response of African savanna ants to long-term fire regimes. Journal of Applied Ecology 41(4): 630-642.
Pietersen, D. W., McKechnie, A. E., & Jansen, R. (2011) A review of the anthropogenic threats faced by Temminck's ground pangolin, Smutsia temminckii, in southern Africa. African Journal of Wildlife Research 44(2): 167-178.
Redford, K. H. (1984) The nutritional value of invertebrates with emphasis on ants and termites as food for mammals. J. Zool Lond 203: 385-395.
Redford, K. H. (1987) Ants and termites as food: patterns of mammalian myrmecophagy. In: Current mammalogy, I. Genoways, H. H. (Ed.). New York: Plenum Press.
Ryder Wilkie, K. T., A. L. Mertl and J. F. A. Traniello. (2010) Species Diversity and Distribution Patterns of the Ants of Amazonian Ecuador. PLoS ONE 5: 131-146.
Samways, M. J. (1983) Community structure of ants (Hymenoptera: Formicidae) in a series of habitats associated with citrus. J. appl. Ecol. 20: 833-847.
Sih, A. (1993) Effects of ecological interactions on forager diets: competition, predation risk, parasitism and prey behaviour. Diet selection: an interdisciplinary approach to foraging behaviour.
Spence, J. R., & Niemelä, J. K. (1994) Sampling carabid assemblages with pitfall traps: the madness and the method. The Canadian Entomologist 126(3): 881-894.
Sudd, J. H. (1982) Ants: foraging, nesting, brood behavior, and polyethism. In Social insects, 4. H. R. Hermann (Ed.). New York: Academic Press.
Sun, N. C. M., Lo, F. H. Y., Chen, B. Y., Yu, H. Y., Liang, C. C., Lin, C. C.,Chen, S. C. , Li, H. F. (2020) Digesta retention time and recovery rates of ants and termites in Chinese pangolins (Manis pentadactyla). Zoo Biology 39(3): 168-175.
Suzuki, A., Thong, S., Tan, S., Iwata, A.(2017) Camera trapping of large mammals in Chhep Wildlife Sanctuary, northern Cambodia. Cambodian Journal of Natural History 2017 (1): 63-75.
Swart, J. M., Richardson, P. R. K., & Ferguson, J. W. H. (1999) Ecological factors affecting the feeding behaviour of pangolins (Manis temminckii). Journal of Zoology 247(3): 281-292.
Van Driesche, R., & Hoddle, M. (2009) Control of pests and weeds by natural enemies: an introduction to biological control. John Wiley and Sons.
Venzon, M., Janssen, A., Pallini, A., & Sabelis, M. W. (2000) Diet of a polyphagous arthropod predator affects refuge seeking of its thrips prey. Animal Behaviour 60(3): 369-375.
Wang, C., J. S. Strazanac and L. Butler. (2001) A Comparison of Pitfall Traps with Bait Traps for Studying Leaf Litter Ant Communities. Entomology Soc 94: 761-765.
Westoby, M. (1978) What are the biological bases of varied diets?. The American Naturalist 112(985): 627-631.
Wu, S. B., Ma, G. Z., Tang, M., Chen, H., & Liu, N. (2002) The status and conservation strategy of pangolin resource in China. Journal of Natural Resources 17(2): 174-180.
Wu, S. B., G. Z. Ma, H. Chen, Z. R. Xu, Y. Y. Li, and N. F. Liu (2004) A preliminary study on burrow ecology of Manis pentadactyla. The Journal of Applied Ecology 15: 401-407.
Wu, S. B., N. F. Liu, Y. Y. Li, and R. Y. Sun (2005) Observation on food habits and foraging behavior of Chinese pangolin (Manis pentadactyla). Chinese Journal of Applied and Environmental Biology 11: 337-341.
Yeatman, E. M., & Greenslade, P. J. M. (1980) Ants as indicators of habitat in three conservation parks in South Australia. South Australian Naturalist 55(2): 20-26.
AntWeb. Version 8.24. California Academy of Sciences, online at https://www.antweb.org Accessed 14 April 2020.
The IUCN Red List of Threatened Species(2019)Version 2018-2, 17 January 2019. http://www.iucnredlist.org.